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Aphid Species File
Species

Cinara (Cinara) pilicornis (Hartig, 1841)

Nomenclature (149)

  • Aphis pilicornis Hartig, 1841: 369.
  • Aphis abietis Walker, 1848: 100.
  • Cinara pilicornis (Hartig, 1841) in Chakrabarti, Medda & Kanturski, 2020: 673.
  • Cinara pilicornis (Hartig, 1841) in Lee, Kanturski, Kim, Glumac & Lee, 2025: 2.

Nomenclature references (92)

  • Albrecht, A.C. (2017) Illustrated identification guide to the Nordic aphids feeding on conifers (Pinophyta) (Insecta, Hemiptera, Sternorhyncha, Aphidomorpha). European Journal of Taxonomy, 338, 1–160. [in English] https://doi.org/10.5852/ejt.2017.338
  • Barbagallo, S. & Stroyan, H.L.G. (1982) Biological, ecological and taxonomic notes on the aphid fauna of Sicily. Frustula Entomologica, Nuova Serie, 3, 1–182. [1980; in English]
  • Binazzi, A. & Scheurer, S. (2009) Atlas of the Honeydew Producing Conifer Aphids of Europe. Aracne editrice S.r.l., Rome, 127 pp. [in English]
  • Binazzi, A. (1978) Contributi alla conoscenza degli afidi delle conifere I. Le specie dei genn. Cinara Curt., Schizolachnus Mordv., Cedrobium Remaud. ed Eulachnus D. Gu. presenti in Italia (Homoptera: Aphidoidea: Lachnidae). Redia, 61, 291–400. [in Italian]
  • Binazzi, A. (1984) Chiave per le specie afidiche più note delle conifere in Europa. Redia, 67(appendix), 547–571. [in Italian]
  • Binazzi, A. (1994) Chiave per le alate virginoparae delle specie del genere Cinara Curtis rinvenute in Italia (Hommoptera Aphididae Lachninae). Redia, 77(2), appendix 1–11. [in Italian]
  • Blanchard, E.E. (1939) Estudio sistemático de los afidoideos argentinos. Physis. Revista de la Sociedad Argentina de Ciencias Naturales, 17, 857–1003.
  • Buckton, G.B. (1881) Monograph of British Aphides. The Ray Society, London, Vol. 3, 142 pp. [in English]
  • Börner, C. (1952) Europae centralis Aphides. Die Blattläuse Mitteleuropas. Namen, Synonyme, Wirtspflanzen, Generationszyklen. In Mitteilungen der Thüringischen Botanischen Gesellschaft: Vol. supplement 3. . . Issue 1, 1–259 pp. [in German]
  • Börner, C.J.B. & Franz, H. (1956) Die Blattläuse des Nordostalpengebietes und sienes Vorlandes. Österreichische Zoologische Zeitschrift, 6(3-5), 297–411. [in German] Available at https://www.zobodat.at/publikation_articles.php?id=59066
  • Börner, C.J.B. & Heinze, K. (1957) Aphidina - Aphidoidea Blattläuse, plantlice (aphids), pucerons (aphides). In L. Reh (Ed.). Handbuch der Pflanzenkrankheiten von Prof. Dr. Paul Sorauer. Paul Parey, Berlin. Vol. 5, Issue 4, 402 pp. [in German]
  • Börner, C.J.B. & Schilder, F.A. (1931) Aphidoidea, Blattläuse. In L. Reh (Ed.). Handbuch der Pflanzenkrankheiten Begründet von Paul Sorauer: Vol. Volume 5. Paul Parey, Berlin. Issue 2, pp. 551–715. [1932; in German]
  • Carrillo, R. (1977) Aphidoidea de Chile II. Agro Sur, 5(2), 109–114.
  • Carter, C.I. & Maslen, N.R. (1982) Conifer lachnids in Britain. Forestry Commission Bulletin, 58, 75 pp. [in English] Available at https://www.forestresearch.gov.uk/publications/archive-conifer-lachnids-in-britain/
  • Carver, M. (1976) New and recent additions to the aphid fauna of Australia (Homoptera: Aphididae). Journal of the Australian Entomological Society, 15, 461–465.
  • Chakrabarti, S., Medda, P.K. & Kanturski, M. (2020) Conifer-feeding aphids (Insecta: Hemiptera: Aphididae) of India, Bhutan and Nepal with descriptions of three new species of the genus Cinara. The European Zoological Journal, 87(1), 659–687. [in English] https://doi.org/10.1080/24750263.2020.1831086
  • Cholodkovsky, N.A. (1892) Zur Kenntnis der Coniferen-Läuse. Zoologischer Anzeiger, 15(384), 66–70. [in German] Available at http://www.biodiversitylibrary.org/page/9740959
  • Cholodkovsky, N.A. (1896) Aphidologische Mittheilungen. Zoologischer Anzeiger, 19(520), 508–513. [in German] Available at http://www.biodiversitylibrary.org/page/9901142
  • Cholodkovsky, N.A. (1896) Zur Kenntnis der auf Fichte (Picea excelsa Lk.) lebenden Lachnus-Arten. Zoologischer Anzeiger, 19(499), 145–150. [in German] Available at http://www.biodiversitylibrary.org/page/9900779
  • Cholodkovsky, N.A. (1898) Explanatory Catalog of the Aphid Collection (Aphidae). Zoological Cabinet of St. Petersburg State University, Saint Petersburg, 24 pp.
  • Cholodkovsky, N.A. (1898) Beiträge zu einer Monographie der Coniferen-Läuse: 2. Die Gattung Lachnus Burm. Proceedings of the Russian Entomological Society, 31, 603–674.
  • Cholodkovsky, N.A. (1902) Second annotated catalogue of aphid (Aphidae) collections in Zoological Cabinet of St. Petersburg Institute of Forestry. Izvestiya S-Peterburgskogo Lesnogo Instituta, 8(2), 1–11.
  • Danielsson, R. (1974) New records of Swedish aphids (Hem. Hom. Aphidoidea). Entomologisk Tidskrift, 95(1), 64–72. [in English] Available at https://www.sef.nu/download/entomologisk_tidskrift/ET%201974/ET1974-64_72.pdf
  • Danielsson, R. (1987) Notes on the taxonomy and nomenclature of some European Cinara species. In Population Structure, Genetics and Taxonomy of Aphids and Thysanoptera. Proceedings of the International Symposium, Smolenice, Czechoslovakia, 9-14 September 1985. SPB Academic Publishing, The Hague, Netherlands. pp. 334–346.
  • Del Guercio, G. (1909) Contribuzione alla conoscenza dei Lacnidi Italiani. Morphologia, sistematica, biologia generale e loro importanza economica. Redia, 5(2), 173–359. [1908; in Italian]
  • Doncaster, J.P. (1961) In Francis Walker's Aphids. British Museum (Natural History), London. 165 pp.
  • Doncaster, J.P. (1973) G.B. Buckton's works on Aphidoidea (Hemiptera). Bulletin of the British Museum (Natural History) Entomology, 28(2), 23–109. [in English]
  • Eastop, V.F. & Hille Ris Lambers, D. (1976) Survey of the World’s Aphids. Dr. W. Junk, The Hague, 573 pp. [in English]
  • Eastop, V.F. (1972) A taxonomic review of the species of Cinara Curtis occurring in Britain (Hemiptera: Aphididae). Bulletin of the British Museum (Natural History) Entomology, 27(2), 103–186. [in English]
  • Eastop, V.F., Miyazaki, M. & Sorin, M. (1998) A check list of Japanese Cinara Curtis (Homoptera: Aphididae) with keys to the species. Miscellaneous Publication of the National Institute of Agro-Environmental Sciences, 22, 1–25. [in English]
  • Fitch, A. (1851) Catalogue with references and descriptions of the insects collected and arranged for the State Cabinet of Natural History. Family Aphidae. Annual Report of the New York State Cabinet of Natural History, Fourth Report, 42–69.
  • Foottit, R.G., Halbert, S.E., Miller, G.L., Maw, H.E.L. & Russell, L.M. (2006) Adventive aphids (Hemiptera: Aphididae) of America north of Mexico. Proceedings of the Entomological Society of Washington, 108(3), 583–610. [in English] Available at https://www.biodiversitylibrary.org/page/30254009
  • Fossel, A. (1970) Anleitung zur Determination einiger in Mitteleuropa verbreiteten Vertreter des Genus Cinara Curt. (Aphidoidea, Lachnidae). 8(5-6), 129–190. [1969]
  • Ghosh, A.K. (1982) The Fauna of India and Adjacent Countries. Homoptera, Aphidoidea. Part 2. Subfamily Lachninae. Zoological Survey of India, Calcutta, 167 pp. [in English]
  • Gittins, A.R., Bishop, G.W., Knowlton, G.F. & Parker, E.J. (1976) An annotated list of the aphids of Idaho (Homoptera: Aphidae). Bulletin of the Idaho Agricultural Experiment Station, 96, 47 pp. [in English]
  • Gutiérrez Alaiz, I., Nieto Nafría, J.M. & Mier Durante, M.P. (1986) Sobre 12 especies de Cinara de España (Hom., Aphidoidea, Lachnidae). 61(1-4), 89–94. [1985]
  • Gómez-Menor Ortega, J. (1962) Lachnidae de España (Hom. Sternorrhyncha). Revista Española de Entomología, 38(3), 347–413. [in Spanish; Castilian]
  • Hartig, T. (1841) Versuch einer Eintheilung der Pflanzenläuse (Phytophthires Burm.) nach der Flügelbildung. Zeitschrift für die Entomologie, 3, 359–376.
  • Havelka, J., Danilov, J. & Rakauskas, R. (2020) Ecological and molecular diversity of Eulachnini aphids (Hemiptera: Aphididae: Lachninae) on coniferous plants in Lithuania. European Journal of Entomology, 117, 199–209. [in English] https://doi.org/10.14411/eje.2020.021
  • Heie, O.E. (1995) The Aphidoidea (Hemiptera) of Fennoscandia and Denmark. VI. Family Aphididae: Part 3 of tribe Macrosiphini of subfamily Aphidinae, and family Lachnidae. Fauna Entomologica Scandinavica, 31, 222 pp. [in English]
  • Heinze, K. (1962) Pflanzenschädliche Blattlausarten der Familien Lachnidae, Adelgidae und Phylloxeridae, eine systematische-faunistische Studie. Deutsche Entomologische Zeitschrift, 9(1-2), 143–227. [in German] https://doi.org/10.1002/mmnd.19620090108
  • Holopainen, J.K. (1983) Conifer aphids (Homoptera: Adelgidae, Lachnidae) from central Finland. 7, 7–12.
  • Hottes, F.C. (1953) Aphidological gleanings. The Pan-Pacific Entomologist, 29(3), 147–155.
  • Hottes, F.C. (1953) Notes on some species of Cinara, with descriptions of two new species from piñon pine (Aphidae). Proceedings of the Biological Society of Washington, 66, 153–158. [in English] Available at http://www.biodiversitylibrary.org/page/36185066
  • Hottes, F.C. (1955) Cinara descriptions (Aphidae). Proceedings of the Biological Society of Washington, 68, 67–77. [in English] Available at http://www.biodiversitylibrary.org/page/34549551
  • Hottes, F.C. (1961) A review and key of North American Cinara (Homoptera: Aphididae) occurring on Picea. Great Basin Naturalist, 21, 35–50. [in English] Available at https://scholarsarchive.byu.edu/gbn/vol21/iss3/1
  • Inouye, M. (1970) Revision of the conifer aphid fauna of Japan (Homoptera, Lachnidae). Bulletin of the Government Forest Expereriment Station, 228, 57–102.
  • Ivanovskaya, O.I. (1977) Families Adelgidae - Chaitophoridae. In Aphids of Western Siberia. Biological Institute, Siberian Branch of the Academy of Sciences of the USSR, Novosibirsk. Vol. 1, 272 pp. [in Russian]
  • Juchnevitch, L.A. (1962) On the aphid fauna of Aphidoidea of softwood in central and south-eastern Kazakhstan. Proceedings of the Institute of Zoology, Academy of Sciences of the Kazakh SSR, 18, 150–154.
  • Kaltenbach, J.H. (1843) Monographie der Familien der Pflanzenläuse (Phytophthires). P. Fagot, Aachen, 222 pp. [in German]
  • Kaltenbach, J.H. (1864) Die deutschen Phytophagen aus der Klasse der Insekten. Verhandlungen des Naturhistorischen Vereines der Preussischen Rheinlande, Westfalens und des Reg.-Bezirks Osnabruck, 21, 228–404.
  • Kirkaldy, G.W. (1906) Catalogue of the genera of the hemipterous family Aphidae, with their typical species, together with a list of the species described as new from 1885 to 1905. The Canadian Entomologist, 38(1), 9–18. [in English] https://doi.org/10.4039/Ent389-1
  • Kloft, W., Kunkel, H. & Ehrhardt, P. (1960) Beitrag zur Lachnidenfauna Mitteleuropas (Homoptera: Aphididae). Beiträge zur Entomologie, 10(1-2), 161–168.
  • Koch, C.L. (1856) Die Pflanzenläuse Aphiden getreu nach dem Leben abgebildet und beschrieben. J.L. Lotzbeck, Nürnberg, Vol. 8, 237–274 pp. [in German]
  • Lampel, G. (1968) Die Biologie des Blattlaus-Generationswechsles Mit besonderer Berücksichtigung terminologischer Aspekte. Gustav Fischer Verlag, Jena, 264 pp.
  • Lee, M., Kanturski, M., Kim, M., Glumac, A. & Lee, S. (2025) First record and integrative analysis of the invasive aphid Cinara pilicornis in South Korea. Scientific Reports, 15(29075), 1–19. [in English] https://doi.org/10.1038/s41598-025-12656-6
  • Mamontova, V.A. (1963) New data on the aphid fauna (Homoptera, Aphidoidea) of Ukraine. Works of the Institute of Zoology, Academy of Sciences of Ukraine RSR, 19, 11–40. [in Ukrainian]
  • Mamontova, V.A. (1964) Aphids of the Ukrainian “Polissie.” Works of the Institute of Zoology, Academy of Sciences of Ukraine RSR, 20, 52–72. [in Ukrainian]
  • Mamontova, V.A. (1972) Aphids - Lachnids. Fauna Ukraini, 20(7), 1–228.
  • Mamontova, V.A. (1991) Remarks on the paper of Yu. M. Rukavishnikov «An attempt of an analysis by karyological method of superspecific taxa in Lachnidae family». Vestnik Zoologii, 1991(4), 83–85.
  • Mamontova, V.A. (2001) Phylogenesis and system of the aphids of the family Lachnidae (Aphidoidea, Homoptera) with taking into account the data concerning caryotypes. Vestnik Zoologii, 35(4), 3–16. [in Russian]
  • Mamontova, V.A. (2008) Evolution, phylogenesis and system of aphids of the lachnid family (Homoptera, Aphidoidea, Lachnidae). Naukova Dumka, Kiev, 207 pp. [in Russian]
  • Mamontova, V.A. (2012) Aphid fauna of the lachnid family (Homoptera, Aphidoidea, Lachnidae) of Eastern Europe and neighboring territories. Naukova Dumka, Kiev, 390 pp. [in Russian]
  • Mordvilko, A.K. (1895) Zur Biologie und Systematik der Baumläuse (Lachninae Pass. partim) des Weichselgebietes. Zoologischer Anzeiger, 18(469), 73–85. [in German] Available at http://www.biodiversitylibrary.org/page/9728416
  • Mordvilko, A.K. (1895) Zur Biologie und Systematik der Baumläuse (Lachninae Pass. partim) des Weichselgebietes. Zoologischer Anzeiger, 18(470), 93–104. [in German] Available at http://www.biodiversitylibrary.org/page/9728436
  • Pashtshenko, N.F. & Lobkova, L.E. (1990) On the fauna of aphids (Homoptera, Aphidinea) of Kamchatka. In A.S. Lelej (Ed.). News of insect systematics of the Soviet Far East. Akademiya Nauk SSSR, Dalnevostochnoe Otdelenie, Vladivostok. pp. 5–27.
  • Pashtshenko, N.F. (1988) Suborder Aphidinea–Aphids. In P.A. Lehr (Ed.). Homoptera and Heteroptera. Keys to the Insects of the Far East of the USSR. Institute of Biology and Soil Sciences, Far East Branch, Academy of Sciences of the USSR, Vladivostok. Vol. 2, pp. 546–686. [in Russian]
  • Patch, E.M. (1912) Aphid pests of Maine. Bulletin of the Maine Agricultural Experiment Station, 202, 159–178. [in English]
  • Pašek, V. (1954) Conifer aphids in Czechoslovakia (Homoptera - Aphidoidea). Slovak Academy of Sciences, Bratislava, 321 pp. [in Czech]
  • Pintera, A. (1966) Revision of the genus Cinara Curt. (Aphidoidea, Lachnidae) in Middle Europe. Acta Entomologica Bohemoslovaca, 63(4), 281–321. [in English]
  • Remaudière, G. & Remaudière, M. (1997) Catalogue of the World’s Aphididae. INRA, Paris, 473 pp. [in French, English]
  • Schouteden, H. (1907) A supplementary list to Kirkaldy's catalogue of the Aphidae described as new from 1885. Annales de la Société Entomologique de Belgique, 50, 30–36. [1906; in English]
  • Smith, C.F. & Cermeli, M. (1979) An annotated list of Aphididae (Homoptera) of the Caribbean Islands and South and Central America. North Carolina Agricultural Experiment Station Technical Bulletin, 259, 131 pp. [in English] Available at https://d.lib.ncsu.edu/collections/catalog/ua101_001_301470_20220527_35188
  • Smith, C.F. & Parron, C.S. (1978) An annotated list of Aphididae (Homoptera) of North America. North Carolina Agricultural Experiment Station Technical Bulletin, 255, 428 pp. [in English] Available at https://d.lib.ncsu.edu/collections/catalog/ua101_001_301466_20220517_34782
  • Steffan, A.W. (1972) Unterordnung Aphidina, Blattläuse. In W. Schwenke (Ed.). Die Forstschädlinge Europas. Ein Hanbuch in fünf Bänden. Erster Band: Würmer, Schnecken, Spinnentiere, Tausendfüssler und hemimetabole Insekten. Paul Parey, Hamburg & Berlin. pp. 162–370. [in German]
  • Stekolshchikov, A.V. & Buga, S.V. (2018) The aphid fauna (Hemiptera, Sternorrhyncha, Aphidomorpha) of Murmansk province (Russia), with description of Aphis khrulevi sp. nov. and Dysaphis karyakini sp. nov. and males of Chaitophorus nigricantis Pintera, 1987. Zootaxa, 4527(4), 451–493.
  • Stekolshchikov, A.V. & Kozlov, M.V. (2012) Distribution records of aphids (Hemiptera: Phylloxeroidea, Aphidoidea) associated with main forest-forming trees in Northern Europe. Entomologica Fennica, 23, 206–214.
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  • Szelegiewicz, H. (1979) Die Verbreitung und Herkunft der Mitteleuropäischen Cinara-Arten (Homoptera, Lachnidae). In Proceedings of the Seventh International Symposium on the Entomofauna of Central Europe. Leningrad. 146–150 pp. [in German]
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  • Zhang, G., Liu, L., He, F. & Zhong, T. (1988) Six new records of Aphidoidea from China. Acta Entomologica Sinica, 31(2), 228.
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Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
TaxonValid names
RankExtantFossilInvalidTotal
Species101920